When lonely, ten-year-old Hal learns that his wealthy but neglectful parents only rented Fleck, the dog he always wanted, he and new friend Pippa take Fleck and four other dogs from the rental agency on a trek from London to Scotland, where Hal's grandparents live.

The journey begins on a ship at sea in 1766, with a boy named Billy Marvel. After surviving a shipwreck, he finds work in a London theatre. There, his family flourishes for generations as brilliant actors until 1900, when young Leontes Marvel is banished from the stage. Nearly a century later, Joseph Jervis runs away from school and seeks refuge with a reclusive uncle in London. Albert Nightingale's strange, beautiful house, with its mysterious portraits...

When his father is arrested as a debtor in 1849 London, fourteen-year-old John Huffman must take on unexpected responsibilities, from asking a distant relative for help to determining why people are spying on him and his family.

After helping prepare their London home for German bombing raids, young Hettie and her brothers are sent to safety in the countryside, where she strives to keep them connected to each other and their parents.